As for the "false eye"-
Who says that an offspring would carry that trait? It would have just as much chance NOT to carry the trait (since one of it parents had it, one didn't)

Genetics is a funny thing- in theory, the way that a genetic strand splits when creating an egg or sperm, 2 children from the same parents could have 0.00% of the same genetic makeup. Literally, 2 brothers could be absolutely, completely, 100% not related to eachother (genetically speaking). Of course the odds of this happening is beyond ludicrous, just from the shear amount of data involved in a DNA sequence. It would be like 2 people constantly flipping coins for a million years, and everytime they came up opposite. That's not gonna happen- but it's possible.

And if we go back to skeletalfrog's paper, it's quite clear that we know less about our beloved pacmans then we think. It states multiple times that there are 2n specimens of ornates as well as 8n's. Therefore, a 2n ornate would be able to breed with a 2n cranwell, but a 2n ornate would not be able to breed with an 8n ornate.

The term "ornate" may be simple symantics.