Update

After a few weeks of research, group discussions, and pictures, I have learned from Corey W that we have Atelopus hoogmoedi and not Atelopus spumarius. "Hoogmoedi" is not a sub-species of A. spumarius but its own species. Quoting Corey: "Spumarius does not range into the Guyana Shield, all the formerly known as spumarius hoogmoedi and spumarius barbitini are now their own species. Just a case of relatives looking similar."

RE: Phylogeography and demography of Guianan harlequin toads (Atelopus): diversification within a refuge - Noonan and Gaucher

Eric