Ummm, you've had these guys 2&1/2 or 3 weeks. I don't think humidity is the problem. But maybe the paper towels. Paper products should be unbleached, as I've heard of fish getting a diease from using cotton in a biologic filter. I think people use the paper to make sure additions are eating(pooping) and to easier gather fecal samples. The dust that comes off white paper towels is much greater than what comes off of brown ones. I've also read red leg looking irritation can come from cardboard or paper towel.
Just my guess.
Try posting a picture of the temporary home





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When she first got Paul he was sleeping deep at the base of the plant so it may have kept him at a little higher humidity so that maybe why he's acting normal. I agree with Ginger about maybe its time to ditch the paper towels. Soil and more plants would allow them to move from wetter to drier areas in the tank as they see fit.
I am 100% sure now that the skin is the problem with little Willie (my nickname to avoid confusion with my other White's). I found him/her this morning looking a very dark orange color, again this is not normal from the day time. S/he hasn't retreated to the plant again and instead decided to stay out in the open for a little while, to my surprise. S/he is still completely uninterested in food, and must be so uncomfortable, I see him/her sitting and climbing awkwardly because of the skin not coming off. 



