also I noticed that u didn't mention any artificial lighting. If u don't have them on a 12 hour cycle, you're stressing him, like they do with a breeding season. I hear there's a 10% mortality rate with breeding, because of the stress through cycling them. It's done by reducing lighting and stop feeding. Then they go in a rain chamber. They don't do this till frogs are 3.
Moral if the story, not lighting them 12 hours causes stress. Get some waxies, and a light. See if that turns him around! Sorry I miss stated before. But without light cycle being regular it does stress them.





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