I have used wingless whenever I used fruit flies. Fruit flies with wings can, if the temperature gets too high mutate to flying flies. And as everyone has said, if one flying fruit fly gets in all bets are off.
I do prefer the wingless because they can not glide so if one manages to miss the tank it falls straight down.
I was going to add this very point. During our unusually hot summer, I purchased a culture online and there was a note in there from them with an appology saying that due to the high temps I may get some able to fly.
I only noticed a couple that could fly so it wasn't a problem.
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"Reverting" to flying is more often caused by mixing or wild ones getting in. I've bred fruit flies for over 30 years and rarely produce fliers.
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